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The game is based in a future where a massive ancient space alien-constructed structure has been discovered buried inside a mountain on an island in the fictional Lingshan Islands, near the coast of the East Philippines. The single-player campaign has the player assume the role of U.S. Army Delta Force soldier Jake Dunn, referred to in-game by his callsign, Nomad. Nomad is armed with various futuristic weapons and equipment, most notably a "Nano Suit" which was inspired by a real-life military concept. In Crysis, the player fights both North Korean and extraterrestrial enemies in various environments on and around the island.
The player can engage enemies in a variety of ways; using stealth or aggression, bullets or non-lethal tranquilizers, ranged rifles or short-range weaponry, and so on. Enemy soldiers employ tactical maneuvers and work as squads. AI soldiers will respond to noise caused by the player, including using signal flares to call for reinforcements. If the player has not been detected in the area, enemies will exhibit relaxed behaviour, but if aware of the player they will draw weapons and become combative.